Ben Adlam Red

I have moved to Google as an AI Resident.


I am a graduate student in applied mathematics at Harvard University. In my research, I apply techniques from probability theory to study the fundamental, mathematical principles of evolution. I focus on macroscopic evolutionary timescales and want to understand how evolution efficiently finds novel functionality. I have also studied the spectra of random matrices. Currently, I work with Martin Nowak and Horng-Tzer Yau. More information can be found in my CV and on my LinkedIn.


  1. Current CRISPR gene drive systems are likely to be highly invasive in wild populations, with Charleston Noble, George M. Church, Kevin M. Esvelt, and Martin A. Nowak. eLife 2018;7:e33423. doi: 10.7554/eLife.33423.
  2. Stationary frequencies and mixing times for neutral drift processes with spatial structure, with Alex McAvoy, Benjamin Allen, and Martin A. Nowak. Proc. R. Soc. A (2018) 474 20180238; doi: 10.1098/rspa.2018.0238.
  3. Spectral statistics of sparse random graphs with a general degree distribution, with Ziliang Che. Preprint. arXiv:1509.03368.
  4. Cellular cooperation with shift updating and repulsion, with Andreas Pavlogiannis, Krishnendu Chatterjee, and Martin A. Nowak. Sci. Rep. 5, 17147 (2015); doi: 10.1038/srep17147.
  5. Amplifiers of selection, with Krishnendu Chatterjee and Martin A. Nowak. Proc. R. Soc. A (2015) 471 20150114; doi: 10.1098/rspa.2015.0114.
  6. Universality of fixation probabilities in randomly structured populations, with Martin A. Nowak. Sci. Rep. 4, 6692 (2014); doi: 10.1038/srep06692
  7. Hitting times of large subsets of ZNK, with Krishnendu Chatterjee and Martin A. Nowak.
  8. The time scale of evolutionary innovation, with Krishnendu Chatterjee, Andreas Pavlogiannis, and Martin A. Nowak. PLoS Comput Biol 10(9): e1003818 (2014). doi:10.1371/journal.pcbi.1003818.


I have served as a teaching fellow for undergraduate and graduate courses in mathematics and computer science. I received the Harvard University Certificate of Distinction in Teaching in Spring 2012, Spring 2015, and Fall 2016. At the Harvard Allston Education Portal, I mentored several students in mathematics, and I am currently a non-residential tutor in mathematics for Harvard's Eliot House.

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As an undergraduate at Harvard, I concentrated in mathematics and wrote my undergraduate thesis on sparse covariance matrices under the supervision of Horng-Tzer Yau.


Phone: +1 (617) 834-5220 Address:
Email: Harvard University, One Brattle Square, suite 6,
Office: 621, Program for Evolutionary Dynamics Cambridge, Massachusetts 02138-3758, USA